1.1 Organisms as Hierarchically Organized Living Systems
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The ultimate goal of my project is to explain how humans evolved a specifi
c kind of vision-related, conscious, creative problem solving ability
I call scenario visualization. However, since conscious creative problem
solving is a psycho-physiological phenomenon that is dependent upon the
workings of the brain and nervous system in the human organism, it is
important for me to give a general philosophical account of organisms and
use this account to explain facts regarding the functioning of the organism’s
systems and processes. I do this in order to offer a philosophy of
biology that is comprehensive enough to account for the levels of biological
phenomena that are relevant to my project. The further upshot is to
lay the groundwork for showing that there is an analogous continuity of
function in the biological world, ranging from the activities of organelles
in a cell to the complex workings of neural networks in a brain from which
conscious abilities emerge (also see Arp, 2005b, 2008a).
In general, biologists and other researchers who describe biological
phenomena are aligned with Mayr (1996, p. 103) in his description of
organisms as “hierarchically organized systems, operating on the basis of
historically acquired programs of information” (Audesirk, Audesirk, &
Beyers, 2002; Gould, 2002; Collier & Hooker, 1999; Eldredge, 1993, 1995;
Bogdan, 1994; Lycan, 1995; Csányi, 1996; Zylstra, 1992; Terzis & Arp,
2008). What exactly is entailed in this description? There are numerous
thinkers who describe organisms and their activities in various ways. In
the next two chapters, I unify several of these conceptions while pointing
to key characteristics of organisms that are relevant to my project as
a whole. In this chapter, I further elucidate the idea that organisms are
hierarchically organized living systems. In the next chapter, after using
ideas and arguments from this chapter in support of certain forms of metaphysical and epistemological forms of emergence, I give further elucidation
to Mayr’s notion that organisms operate on the basis of historically
acquired programs of information, as well as ratify Plotkin’s claim that
biological phenomena only make complete sense in light of evolutionary
theory, by endorsing a hybrid view of functions based in both the Cummins
organizational and the Griffi ths/Godfrey-Smith modern history accounts.
According to Mayr (1996), an organism is a hierarchically organized
living system. What exactly does this mean? We can defi ne an organism as
a hierarchically organized living system made up of components that are
engaged in processes constituting coordinated subsystems, with the product
of these processes and subsystems being homeostasis relative to their operations,
producing the overall homeostasis of the organism. As a system, an
organism is a unifi ed entity that is explainable in terms of the properties
of its components, the interactions of these components, and the overall
coordination of these components. As a living system, an organism has to
be made up of at least one cell, the basic unit of life. To understand what
it means for an organism to be a hierarchically organized living system, we
need to investigate the properties of the components of this kind of system.
These properties include what I call (1) internal–hierarchical data exchange,
(2) data selectivity, (3) informational integration, and (4) environmental–
organismic information exchange (also Arp, 2005b, 2008a). When I describe
each of these properties, the interactions of the components of this kind
of living system, as well as the overall coordination of these components,
will become evident.
However, before investigating the fi rst of these properties in an organism,
namely, internal–hierarchical data exchange, it is necessary to explicate
the words component and homeostasis utilized in the above defi nition
of an organism. The word component is a term that can be used analogously
to refer to either a part of a process, a part of a subsystem, or a part of a
system. In the most general of terms, an organism is a unifi ed living system
made up of subsystems. In turn, these subsystems are made up of processes,
and these processes are the activities in which the components are engaged.
The components of an organism range from the organelles performing
processes in a cell, to cells performing processes in an organ, to organs
performing processes in a subsystem, to subsystems performing processes
in the whole system itself, that is, the organism. Thus, for example, the
respiratory subsystem works with other subsystems in an organism like a
dog to maintain its life: the respiratory subsystem would be considered as
one component of the entire dog, envisioned as one whole system; the
lung would be considered as one component of the respiratory subsystem of the dog; lung cellular tissue comprising one of the lobes of its lung
would be considered as one component of the lung; and the particular
kind of cell that comprises lung tissue is made up of organelles, the basic
components of cells.
Homeostasis refers to the relatively constant or stable coordination of
functioning among the components in the organismic hierarchy, given
the interaction of these components with environmental pressures internal
to and external to the organism. I will have more to say about internal
versus external environmental pressures later in this chapter. For now,
suffi ce it to say that there are environments exerting pressures upon the
subsystems and processes internal to an organism, as well as environments
exerting pressures upon the organism as a whole that are external to it.
The components that make up an organism, as well as the organism itself,
are able to respond effectively to the ever-changing environmental pressures
by adjusting and readjusting their activities so as to continue their
respective operations with a degree of stability. When a subsystem or
process in an organism is operating with a degree of stability despite environmental
pressures—for example, when the cell wall actually performs
the activity of allowing nutrients into the cell, or when a heart actually
performs the activity of pumping blood, or when the body of an animal
actually cools itself through perspiration because its temperature has been
raised above a certain degree—it is said to be functioning properly. I will
have more to say about functions in the next chapter.
We can draw a distinction between what I will call particularized homeostasis
and generalized homeostasis. Particularized homeostasis refers to the
end product of the proper functioning of the particular processes and subsystems
in an organism being the relatively constant coordination among
the components that make up the processes and subsystems, given environmental
pressures that are internal to the organism. Generalized homeostasis
refers to the overall maintenance of the life of an organism being the
result of the proper functioning of the processes and subsystems, given
environmental pressures that are external to the organism. The overall
homeostasis of the living system is maintained because homeostasis is
maintained at the levels of the subsystems and processes.
If the various processes and subsystems of an organism are functioning
properly in their internal environments—thereby producing particularized
homeostasis—the organism is able to live its life effectively in some
external environment. This proper functioning that yields internal homeostasis
takes place at levels in the hierarchy of the organism ranging from
the coordinated activities of organelles in the cell, to cells performing coordinated processes in an organ, to organs performing coordinated processes
in a subsystem, to subsystems performing coordinated activities in
an organism. Thus, in reference to our example of the dog: the dog is able
to live its life in some external environment precisely because of the
overall relatively constant coordination of the subsystems in its body; in
turn, a particular subsystem, like the respiratory subsystem, functions
properly because of the relatively constant coordination of cellular processes;
and the cells themselves function properly because of the relatively
constant coordination among the various organelles.
The subsystems and processes of an organism can be understood as
functioning at various levels of operation, from lower levels to higher
levels. The determination of a subsystem as existing at a certain level
depends upon the way in which the processes of the subsystem operate
and, in turn, the way in which the subsystems operate in the organism as
a whole. Lower level processes operate in certain ways and form the basis
for higher level processes and subsystems. In turn, higher level subsystems
and processes are comprised of lower level processes and utilize the information
from these lower levels to perform their own operations. In this
sense, along with Audesirk et al. (2002), Lycan (1995), and Salthe &
Matsuno (1995), we could say that higher level subsystems are the phenomena
that literally emerge from lower level subsystems and processes.
Later in this chapter and the next, I will have more to say about emergence
as well as about higher levels exhibiting control over lower levels—in terms
of higher levels selecting and integrating information from lower levels—in
an organismic hierarchy.
The organism can be conceptualized as a hierarchical organization
whereby levels of operation, in the forms of subsystems and processes,
function interdependently with one another in this unifi ed system. A
schematization of this hierarchical system is shown in fi gure 1.1. The
organism is represented by the large partitioned triangle that contains the
smaller partitioned triangles within it; the biggest triangles within the one
large triangle represent subsystems, the smaller triangles within those subsystems
represent processes, the smallest triangles within those processes
represent components of processes, and the partitions represent levels of
operation. Some of the triangles overlap, signifying that the subsystems
are interdependently related to one another. For example, in a hierarchically
ordered system like the mammal, the nervous (sub)system is dependent
upon the respiratory and circulatory (sub)systems, primarily for a
process of oxygen transfer to the nerve cells and brain cells of the nervous
(sub)system. At the same time, the processes of the respiratory and circulaOrganisms tory (sub)systems are dependent upon the processes of the nervous
(sub)system—found, specifi cally, in the medulla of the brain—for their
activities.
The ultimate goal of my project is to explain how humans evolved a specifi
c kind of vision-related, conscious, creative problem solving ability
I call scenario visualization. However, since conscious creative problem
solving is a psycho-physiological phenomenon that is dependent upon the
workings of the brain and nervous system in the human organism, it is
important for me to give a general philosophical account of organisms and
use this account to explain facts regarding the functioning of the organism’s
systems and processes. I do this in order to offer a philosophy of
biology that is comprehensive enough to account for the levels of biological
phenomena that are relevant to my project. The further upshot is to
lay the groundwork for showing that there is an analogous continuity of
function in the biological world, ranging from the activities of organelles
in a cell to the complex workings of neural networks in a brain from which
conscious abilities emerge (also see Arp, 2005b, 2008a).
In general, biologists and other researchers who describe biological
phenomena are aligned with Mayr (1996, p. 103) in his description of
organisms as “hierarchically organized systems, operating on the basis of
historically acquired programs of information” (Audesirk, Audesirk, &
Beyers, 2002; Gould, 2002; Collier & Hooker, 1999; Eldredge, 1993, 1995;
Bogdan, 1994; Lycan, 1995; Csányi, 1996; Zylstra, 1992; Terzis & Arp,
2008). What exactly is entailed in this description? There are numerous
thinkers who describe organisms and their activities in various ways. In
the next two chapters, I unify several of these conceptions while pointing
to key characteristics of organisms that are relevant to my project as
a whole. In this chapter, I further elucidate the idea that organisms are
hierarchically organized living systems. In the next chapter, after using
ideas and arguments from this chapter in support of certain forms of metaphysical and epistemological forms of emergence, I give further elucidation
to Mayr’s notion that organisms operate on the basis of historically
acquired programs of information, as well as ratify Plotkin’s claim that
biological phenomena only make complete sense in light of evolutionary
theory, by endorsing a hybrid view of functions based in both the Cummins
organizational and the Griffi ths/Godfrey-Smith modern history accounts.
According to Mayr (1996), an organism is a hierarchically organized
living system. What exactly does this mean? We can defi ne an organism as
a hierarchically organized living system made up of components that are
engaged in processes constituting coordinated subsystems, with the product
of these processes and subsystems being homeostasis relative to their operations,
producing the overall homeostasis of the organism. As a system, an
organism is a unifi ed entity that is explainable in terms of the properties
of its components, the interactions of these components, and the overall
coordination of these components. As a living system, an organism has to
be made up of at least one cell, the basic unit of life. To understand what
it means for an organism to be a hierarchically organized living system, we
need to investigate the properties of the components of this kind of system.
These properties include what I call (1) internal–hierarchical data exchange,
(2) data selectivity, (3) informational integration, and (4) environmental–
organismic information exchange (also Arp, 2005b, 2008a). When I describe
each of these properties, the interactions of the components of this kind
of living system, as well as the overall coordination of these components,
will become evident.
However, before investigating the fi rst of these properties in an organism,
namely, internal–hierarchical data exchange, it is necessary to explicate
the words component and homeostasis utilized in the above defi nition
of an organism. The word component is a term that can be used analogously
to refer to either a part of a process, a part of a subsystem, or a part of a
system. In the most general of terms, an organism is a unifi ed living system
made up of subsystems. In turn, these subsystems are made up of processes,
and these processes are the activities in which the components are engaged.
The components of an organism range from the organelles performing
processes in a cell, to cells performing processes in an organ, to organs
performing processes in a subsystem, to subsystems performing processes
in the whole system itself, that is, the organism. Thus, for example, the
respiratory subsystem works with other subsystems in an organism like a
dog to maintain its life: the respiratory subsystem would be considered as
one component of the entire dog, envisioned as one whole system; the
lung would be considered as one component of the respiratory subsystem of the dog; lung cellular tissue comprising one of the lobes of its lung
would be considered as one component of the lung; and the particular
kind of cell that comprises lung tissue is made up of organelles, the basic
components of cells.
Homeostasis refers to the relatively constant or stable coordination of
functioning among the components in the organismic hierarchy, given
the interaction of these components with environmental pressures internal
to and external to the organism. I will have more to say about internal
versus external environmental pressures later in this chapter. For now,
suffi ce it to say that there are environments exerting pressures upon the
subsystems and processes internal to an organism, as well as environments
exerting pressures upon the organism as a whole that are external to it.
The components that make up an organism, as well as the organism itself,
are able to respond effectively to the ever-changing environmental pressures
by adjusting and readjusting their activities so as to continue their
respective operations with a degree of stability. When a subsystem or
process in an organism is operating with a degree of stability despite environmental
pressures—for example, when the cell wall actually performs
the activity of allowing nutrients into the cell, or when a heart actually
performs the activity of pumping blood, or when the body of an animal
actually cools itself through perspiration because its temperature has been
raised above a certain degree—it is said to be functioning properly. I will
have more to say about functions in the next chapter.
We can draw a distinction between what I will call particularized homeostasis
and generalized homeostasis. Particularized homeostasis refers to the
end product of the proper functioning of the particular processes and subsystems
in an organism being the relatively constant coordination among
the components that make up the processes and subsystems, given environmental
pressures that are internal to the organism. Generalized homeostasis
refers to the overall maintenance of the life of an organism being the
result of the proper functioning of the processes and subsystems, given
environmental pressures that are external to the organism. The overall
homeostasis of the living system is maintained because homeostasis is
maintained at the levels of the subsystems and processes.
If the various processes and subsystems of an organism are functioning
properly in their internal environments—thereby producing particularized
homeostasis—the organism is able to live its life effectively in some
external environment. This proper functioning that yields internal homeostasis
takes place at levels in the hierarchy of the organism ranging from
the coordinated activities of organelles in the cell, to cells performing coordinated processes in an organ, to organs performing coordinated processes
in a subsystem, to subsystems performing coordinated activities in
an organism. Thus, in reference to our example of the dog: the dog is able
to live its life in some external environment precisely because of the
overall relatively constant coordination of the subsystems in its body; in
turn, a particular subsystem, like the respiratory subsystem, functions
properly because of the relatively constant coordination of cellular processes;
and the cells themselves function properly because of the relatively
constant coordination among the various organelles.
The subsystems and processes of an organism can be understood as
functioning at various levels of operation, from lower levels to higher
levels. The determination of a subsystem as existing at a certain level
depends upon the way in which the processes of the subsystem operate
and, in turn, the way in which the subsystems operate in the organism as
a whole. Lower level processes operate in certain ways and form the basis
for higher level processes and subsystems. In turn, higher level subsystems
and processes are comprised of lower level processes and utilize the information
from these lower levels to perform their own operations. In this
sense, along with Audesirk et al. (2002), Lycan (1995), and Salthe &
Matsuno (1995), we could say that higher level subsystems are the phenomena
that literally emerge from lower level subsystems and processes.
Later in this chapter and the next, I will have more to say about emergence
as well as about higher levels exhibiting control over lower levels—in terms
of higher levels selecting and integrating information from lower levels—in
an organismic hierarchy.
The organism can be conceptualized as a hierarchical organization
whereby levels of operation, in the forms of subsystems and processes,
function interdependently with one another in this unifi ed system. A
schematization of this hierarchical system is shown in fi gure 1.1. The
organism is represented by the large partitioned triangle that contains the
smaller partitioned triangles within it; the biggest triangles within the one
large triangle represent subsystems, the smaller triangles within those subsystems
represent processes, the smallest triangles within those processes
represent components of processes, and the partitions represent levels of
operation. Some of the triangles overlap, signifying that the subsystems
are interdependently related to one another. For example, in a hierarchically
ordered system like the mammal, the nervous (sub)system is dependent
upon the respiratory and circulatory (sub)systems, primarily for a
process of oxygen transfer to the nerve cells and brain cells of the nervous
(sub)system. At the same time, the processes of the respiratory and circulaOrganisms tory (sub)systems are dependent upon the processes of the nervous
(sub)system—found, specifi cally, in the medulla of the brain—for their
activities.